Given the question: Given the following passage  "Mahayana Buddhism flourished in India from the 5th century CE onwards, during the dynasty of the Guptas. Mahāyāna centres of learning were established, the most important one being the Nālandā University in north-eastern India.",  answer the following question. Note that the answer is present within the text.  Question: The centres of learning were all where?
The answer is:
Mahāyāna centres of learning were established, the most important one being the Nālandā University in north-eastern India