Given the question: Question: "The centres of learning were all where?"  Context: "Mahayana Buddhism flourished in India from the 5th century CE onwards, during the dynasty of the Guptas. Mahāyāna centres of learning were established, the most important one being the Nālandā University in north-eastern India."  Answer:
The answer is:
Mahāyāna centres of learning were established, the most important one being the Nālandā University in north-eastern India